(photographed in Selangor, Malaysia)

Shih, H.-T., Ng, P. K. L., Davie, P. J. F., Schubart, C. D., Türkay, M., Naderloo, R., Jones, D. & Liu, M.-Y. 2016. Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera. Raffles Bulletin of Zoology 64: 139-175.

Shih, H.-T., Ng, P. K. L. & Liu, M.-Y., 2013. Systematics of the Indo-West Pacific broad-fronted fiddler crabs (Crustacea: Ocypodidae: genus Uca). The Raffles Bulletin of Zoology 61(2): 641-649.